Prof. Maria Włodarska-Kowalczuk
Department of Marine Ecology
Institute of Oceanology, Polish Academy of Sciences
Powstańcow Warszawy 55
81-712 Sopot
tel: (+ 48 58) 731 17 78
fax: (+ 48 58) 551 21 30
e-mail: maria [at]

  • 2018 - Professor (full professor rank)
  • 2018 - present: head of the Department of Marine Ecology
  • 2006 - present: head of the Benthic Ecology Laboratory at IO PAN
  • 2009 - present: assistant professor at the IO PAN, Department of Marine Ecology
  • 2009: habilitation at IO PAN
  • 2001: PhD at University of Gdańsk, Faculty of Biological Oceanography
  • 1999 - 2009: research scientist at IO PAN, Department of Marine Ecology
  • 1995: MSc at University of Gdańsk, Faculty of Biological Oceanography
  • Role of ecosystem engineers (kelp beds, macrophytes in seagrass meadows, bioturbators) in functioning of the benthic systems. (link to Hornsund 2003 fieldwork report)
  • Environmental drivers of patterns of benthic diversity, including both taxonomic and functional diversity, food web complexity and size structure of benthic communities.
  • Benthic Biomass Size Spectra (BBSS) as descriptors of community diversity and function.
  • Arctic Molluscs (link to Kongsfjorden Mollusca species picture gallery)
  • 2018-2020 ABeFun - Arctic benthos functioning response to climate warming induced changes in warm water advection and food supply (Polish National Science Center, coordinator: dr Maria Włodarska-Kowalczuk)
  • 2014-2017 DWARF - Declining size - a general response to climate warming in Arctic fauna? (Polish-Norwegian Research Programme, coordinator: dr Maria Włodarska-Kowalczuk)
  • 2014-2017 POLNOR - The Changing Ocean of the Polar North (Polish-Norwegian Research Programme, coordinator: dr Piotr Kukliński)
  • 2014-2017 GLAERE - Glaciers as Arctic Ecosystem Refugia (Polish-Norwegian Research Programme, coordinator: prof. Jan Marcin Węsławski)
  • 2013-2016 BIOSIZE - Biomass Size Spectra and secondary production - benthic response to environmental variability in Arctic coastal waters, shelf and deep-sea (Polish National Science Center, coordinator: dr Maria Włodarska-Kowalczuk)
  • 2012 -2016 GAME – Growing of the Arctic Marine Ecosystem (Polish National Scienec Center – coordinator: prof. Jan Marcin Węsławski)
  • 2013-2016 4AGE - Does low taxonomic diversity determine trophic structure impoverishment in benthic communities in Arctic fjords? (Polish National Science Center, coordinator: dr Joanna Legeżyńska)

Włodarska-Kowalczuk M., Pawłowska J., Zajączkowski M., 2013, Do foraminifera mirror diversity and distribution patterns of macrobenthic fauna in an Arctic glacial fjord?, Marine Micropaleontology 103, 30-39

Włodarska-Kowalczuk M., Renaud P.E., Węsławski J.M., Cochrane S.K.J., Denisenko S.G., 2012, Species diversity, functional complexity and rarity in arctic fjordic versus open shelf benthic system. Marine Ecology Progress Series 463:73-87

Włodarska-Kowalczuk, M., Kuklinski, P., Ronowicz, M., Legezynska, J., Gromisz, S. 2009. Assessing species richness of macrofauna associated with macroalgae in Arctic kelp forests (Hornsund, Svalbard). Polar Biology 32, 897-905

Włodarska-Kowalczuk M., Węsławski, J.M., 2008, Mesoscale spatial structures of soft-bottom macrozoobenthos: physically controlled/impoverished versus biologically accommodated/diverse communities, Mar Ecol Prog Ser 356, 215-224

Włodarska-Kowalczuk M., Kedra M., 2007, Surrogacy in natural patterns of benthic distribution and diversity: selected taxa versus lower taxonomic resolution, Mar Ecol Prog Ser , 351, 53-63


And here is link to my Research Gate

  • Balazy P, Anderson MJ, Chelchowski M, Włodarska-Kowalczuk M, Kuklinski P, Berge J, 2021, Shallow-Water Scavengers of Polar Night and Day – An Arctic Time-Lapse Photography Study. Front. Mar. Sci. 8:656772
  • Mazurkiewicz M, Meyer-Kaiser K, Sweetman AK, Renaud PE, Włodarska–Kowalczuk M, 2021, Megabenthic standing stocks and organic carbon demand in a warming Arctic, Progress in Oceanography, 196, 102616
  • Kotwicki, L., Węsławski, J.M., Włodarska-Kowalczuk, M., Mazurkiewicz, M., Wenne, R., Zbawicka, M., Minchin, D., Olenin, S., 2021, The re-appearance of the Mytilus spp. complex in Svalbard, Arctic, during the Holocene: The case for an arrival by anthropogenic flotsam, Global and Planetary Change 202, 103502
  • Węsławski, J.M, Legeżyńska, J., Włodarska-Kowalczuk, M., 2020, Will shrinking body size and increasing species diversity of crustaceans follow the warming of the Arctic littoral? Ecology and Evolution 10, 10305-10313.
  • Górska, B., Soltwedel, T., Schewe, I., Włodarska-Kowalczuk, M., 2020, Bathymetric trends in biomass size spectra, carbon demand, and production of Arctic benthos (76-5561 m, Fram Strait), Progress in Oceanography 186.
  • Mazurkiewicz, M., Górska, B., Renaud, P.E. & Włodarska-Kowalczuk, M., 2020, Latitudinal consistency of biomass size spectra - benthic resilience despite environmental, taxonomic and functional trait variability. Scientific Reports, 10, 4164.
  • Ronowicz, M., Włodarska-Kowalczuk, M., Kuklinski, P., 2020, Glacial and depth influence on sublittoral macroalgal standing stock in a high-Arctic fjord, Continental Shelf Research 194.
  • Zaborska A., Włodarska-Kowalczuk M., Legeżyńska J., Jankowska E., Winogradow A., Deja K., 2018, Sedimentary organic matter sources, benthic consumption and burial in west Spitsbergen fjords – Signs of maturing of Arctic fjordic systems?, Journal of Marine Systems 180: 112-123.
  • Degen R., Aune M., Bluhm B.A., Cassidy C., Kędra M., Kraan C., Vandepitte L., Włodarska-Kowalczuk M., Zhulay I., Albano P.G., Bremner J., Grebmeier J.M., Link H., Morata N., Nordström M.C., Shojaei G.M., Sutton L., Zuschin M., 2018, Trait-based approaches in rapidly changing ecosystems: A roadmap to the future polar oceans, Ecological Indicators 91: 722-736.
  • Zawierucha, K., Podkowa, P., Marciniak, M., Gąsiorek, P., Zmudczyńska-Skarbek, K., Janko, K., Włodarska-Kowalczuk, M., 2018, Temperature (latitude) and nutrient (seabird guano) effects on limno-terrestrial Tardigrada (Testechiniscus spitsbergensis and Pilatobius recamieri) body size, Polar Research 37:1.
  • Ronowicz, M., Kukliński, P., Włodarska-Kowalczuk, M., 2018, Diversity of kelp holdfast-associated fauna in an Arctic fjord - Inconsistent responses to glacial mineral sedimentation across different taxa, Estuarine and Coastal Shelf Science 205, 100-109.
  • Jankowska, E., De Troch, M., Michel, L.N., Lepoint, G., Włodarska-Kowalczuka, M., 2018, Modification of benthic food web structure by recovering seagrass meadows, as revealed by trophic markers and mixing models,Ecological Indicators 90: 28-37 .
  • Stępień A., Kukliński P., Włodarska-Kowalczuk M., Krzemińska M., Gudmundsson, 2017, Bryozoan zooid size and shape variability across bathymetric gradient. Marine Biology 164(10): 197.
  • Drewnik, A., Węsławski J.M., Włodarska-Kowalczuk M., 2017, Benthic Crustacea and Mollusca distribution in Arctic fjord–case study of patterns in Hornsund, Svalbard, Oceanologia 59, 565-575.
  • Górska B., Włodarska-Kowalczuk M, Food and disturbance effects on Arctic benthic biomass and production size spectra. Progress in Oceanography (2017), 152: 50-61.
  • Legeżyńska J., M. Włodarska-Kowalczuk, M. Głuchowska, M. Ormańczyk, M. Kędra, J.M. Węsławski, 2017, The malacostracan fauna of two Arctic fjords (west Spitsbergen): the diversity and distribution patterns of its pelagic and benthic components. Oceanologia 59: 541-564
  • Mazurkiewicz, M., Gromisz S., Legeżyńska J., Włodarska-Kowalczuk M., 2017, First record of Lacydonia eliasoni Hartmann-Schröder, 1996 (Polychaeta: Phyllodocida) in the European Arctic. Polish Polar Research 38(2): 173-183.
  • Węsławski, J.M., Urbański, J., Głuchowska, M., Grzelak, K., Kotwicki, L., Kwaśniewski, S., Legeżyńska, J., Wiktor, J., Włodarska-Kowalczuk, M., Zaborska, A., Zajączkowski, M., Stempniewicz, L. 2017. Can seabirds modify carbon burial in fjords? Oceanologia 59, 603-611.
  • Zaborska A., Beszczyńska-Moller A., Włodarska-Kowalczuk M., 2017, History of heavy metal accumulation in the Svalbard area: distribution, origin and transport pathways. Environmental Pollution 231: 437-450
  • Włodarska-Kowalczuk M, Górska B., Deja K, Morata N, Do benthic meio- and macrofaunal communities respond to seasonality in pelagial processes in an Arctic fjord (Kongsfjorden, Spitsbergen)? Polar Biology, 2016, 39(11): 2115-2129
  • Jankowska E., Löic N. Michel, Zaborska A, Włodarska-Kowalczuk M, Sediment carbon sink in low density temperate eelgrass meadows (Baltic Sea), Journal of Geophysical Research: Biogeosciences, (2016), 121, 2918-2934.
  • Mazurkiewicz M, Górska B., Jankowska E, Włodarska-Kowalczuk M, Assessment of nematode biomass in marine sediments - semi-automated image analysis method. Limnology and Oceanography: Methods, 2016, 14(12): 816-827
  • Włodarska-Kowalczuk M., Mazurkiewicz M., Jankowska E., Kotwicki L., Damrat M., Zajączkowski M., 2016, Effects of fluvial discharges on meiobenthic and macrobenthic variability in the Vistula River prodelta (Baltic Sea), Journal of Marine Systems 157, 135-146
  • Balazy P, Kuklinski P, Włodarska-Kowalczuk M, Gluchowska M, Barnes D. K. A. 2016. Factors affecting biodiversity on hermit crab shells; Hydrobiologia 773 (1), 207-224.
  • Deja K, Węsławski JM, Borszcz T, Włodarska-Kowalczuk M, Kukliński P, Balazy P, Kwiatkowska P (2016) Recent distribution of Echinodermata species in Spitsbergen coastal waters. Polish Polar Research 37(4), 511-526
  • Drewnik, A., Węsławski, J.M., Włodarska-Kowalczuk, M., Łacka M., Prominska, A., Głuchowska M., Zaborska, A. 2016 From the worm’s point of view. I: Environmental settings of benthic ecosystems in Arctic fjord (Hornsund, Spitsbergen) Polar Biology 39, 1411–1424
  • Soltwedel., T., Bauerfeind, E., Bergmann, M., Bracher, A., Budaeva, N., Busch, K., Cherkasheva, A., Fahl, K., Grzelak, K., Hasemann, C., Jacob, M., Kraft, A., Lalande, C., Metfis, K., Nöthig, M-E., Meyer, K., Quéric, N-V., Schewe, I., Włodarska-Kowalczuk, M., Klages, M. 2016. Natural variability or anthropogenically-induced variation? Insights from 15 years of multidisciplinary observations at the arctic marine LTER site HAUSGARTEN. Ecological Indicators 65, 89-102
  • Bałazy P, Kukliński P, Włodarska-Kowalczuk M., Barnes DKA., Kedra M., Legezyńska J., Węsławski J.M., 2015, Hermit crabs (Pagurus spp.) at their northernmost range: distribution, abundance and shell use in the European Arctic. Polar Research, 34, 21412
  • Berge J., Daase M., Renaud P.E., Ambrose W.A., Jr., Darnis G., Last K.S., Leu E., Cohen J.H., Johnsen G., Moline M.A., Cottier F., Varpe Ø., Shunatova N., Bałazy P., Morata N., Massabuau J-Ch., Falk-Petersen S., Kosobokova K., Hoppe C.J.M., Węsławski J.M., Kukliński P., Legezyńska J.,., Nikishina D., Cusa M., Kedra M., Włodarska_Kowalczuk M.,., Vogedes D., Camus L., Tran D., Michaud E., Gabrielsen T.M., Granovitch A., Gonchar A., Krapp R. and Callesen T.A. 2015 Unexpected levels of biological activity during the polar night offer new perspectives on a warming Arctic, Current Biology, 2015, 25, 1–7
  • Jankowska E, Jankowska K, Włodarska-Kowalczuk M.,2015, Seagrass vegetation and meiofauna enhance the bacterial abundance in the Baltic Sea sediments (Puck Bay). Environmental Science and Pollution Research.22: 14372-14378
  • Legeżyńska J., Deja K., Włodarska-Kowalczuk M.,2015, First record of the family Dexaminidae and species Guernea (Prinassus) nordenskioldi in the west Spitsbergen fjords (Svalbard, Greenland Sea) Marine Biodiversity Records 8
  • Morata, N., Michaud, E., Włodarska-Kowalczuk, M., 2015, Impact of early food input on the Arctic benthos activities during the polar night. Polar Biology 38, 99-114
  • Włodarska-Kowalczuk M, Jankowska E, Kotwicki L, Balazy P,2014, Evidence of Season-Dependency in Vegetation Effects on Macrofauna in Temperate Seagrass Meadows (Baltic Sea). PLoS ONE 9(7): e100788.
  • Górska B, Grzelak K, Kotwicki L, Hasemann C, Schewe I, Soltwedel T, Włodarska-Kowalczuk M, 2014, Bathymetric variations in vertical distribution patterns of meiofauna in the surface sediments of the deep Arctic ocean (HAUSGARTEN, Fram strait), Deep-Sea Research 91, 36–49
  • Jankowska E., Włodarska-Kowalczuk M., Kotwicki L., Balazy P., Kuliński K., 2014, Seasonality in vegetation biometrics and its effects on sediment characteristics and meiofauna in Baltic seagrass meadows, Estuarine Coastal and Shelf Science 139, 159-170
  • Włodarska-Kowalczuk M, Balazy P, Wiktor J, Kobos J, Zajączkowski M, Moskal W, 2014, Large red cyanobacterial mats (Spirulina subsalsa Oersted ex Gomont) in shallow sublittoral of the Southern Baltic. Oceanologia 56 (3), 661–666
  • Włodarska-Kowalczuk M., Pawłowska J., Zajączkowski M., 2013, Do foraminifera mirror diversity and distribution patterns of macrobenthic fauna in an Arctic glacial fjord?, Marine Micropaleontology 103, 30-39
  • Ronowicz M., Włodarska-Kowalczuk M., Kuklinski P., 2013, Depth- and substrate-related patterns of species richness and distribution of hydroids (Cnidaria, Hydrozoa) in Arctic coastal waters (Svalbard), Marine Ecology - an Evolutionary Perspective, 34 (1) SI,165-176
  • Ronowicz M, Legeżyńska J, Kukliński P, Włodarska-Kowalczuk M., 2013, Kelp forest as a habitat for mobile epifauna, case study of Caprella septentrionalis Kröyer, 1838 in an Arctic glacial fjord, Polar Research, , 32, 1-6
  • Ronowicz M., Włodarska-Kowalczuk M., Kuklinski P., 2013, Hydroid epifaunal communities in Arctic coastal waters (Svalbard): effects of substrate characteristics, Polar Biology 36 (5), 705-718
  • Węsławski J.M., Kryla-Straszewska L., Warzocha J., Urbanski J., Włodarska-Kowalczuk M., Kotwicki L., 2013, How lonely they are? A degree of isolation among macrozoobenthos species in the Marine Protected Area, the Bay of Puck, the Southern Baltic, Oceanological and Hydrobiological Studies 42 (3), 289-295
  • Węsławski J.M., Kryla-Straszewska L., Piwowarczyk J., Urbanski J., Warzocha J., Kotwicki L., Włodarska-Kowalczuk M., Wiktor J., 2013, Habitat modelling limitations - Puck Bay, Baltic Sea - a case study, Oceanologia 55 (1), 167-183
  • Josefson, A., Mokievski, V., Bergmann B., Blicher M.E., Bluhm B., Cochrane S., Denisenko N.V., Hasemann C., Jørgensen L.L., Klages M., Schewe I., Sejr M.K., Soltwedel T., Węsławski J.M., Włodarska-Kowalczuk M., 2013, „Marine Invertebrates” in: Meltofte H. (ed.) Arctic Biodiversity Assessment. Status and trends in Arctic biodiversity. Conservation of Arctic Flora and Fauna, Arctic Council
  • Węsławski, J.M, Włodarska-Kowalczuk, M., Kędra, M., Legeżyńska, J., Kotwicki, L., 2012, Eight species that rule today's European Arctic fjord benthos, Polish Polar Research 33, 225-238
  • Włodarska-Kowalczuk M., Renaud P.E., Węsławski J.M., Cochrane S.K.J., Denisenko S.G., 2012, Species diversity, functional complexity and rarity in arctic fjordic versus open shelf benthic system. Marine Ecology Progress Series 463:73-87
  • Bluhm BA, Ambrose Jr WG, Bergmann M, Clough LM, Gebruk AV, Hasemann C, Iken K, Klages M, MacDonald IR, Renaud PE, Schewe I, Soltwedel T, Włodarska-Kowalczuk M, 2011, Diversity of the Arctic deep-sea benthos, Marine Biodiversity 41, 87-107
  • Pawłowska J., Włodarska-Kowalczuk M., Zajączkowski M., Nygård H., Berge J. 2011. Seasonal variability of meio- and macrobenthic standing stocks and diversity in an Arctic fjord (Adventfjorden, Spitsbergen). Polar Biology 34: 833-845
  • Piepenburg D, Archambault P, Ambrose Jr WG, Blanchard A, Bluhm BA, Carroll ML, Conlan KE, Cusson M, Feder HM, Grebmeier JM, Jewett SC, Lévesque M, Petryashev VV, Sejr MK, Sirenko BI, Włodarska-Kowalczuk M, 2011, Towards a pan-Arctic inventory of the species diversity of the macro- and megabenthic fauna of the Arctic shelf seas, Marine Biodiversity 41, 51-70
  • Ronowicz M., Włodarska-Kowalczuk, M., Kukliński, P., 2011, Patterns of hydroid (Cnidaria, Hydrozoa) species richness and distribution in an Arctic glaciated fjord, Marine Ecology Progress Series 463:73-87
  • Wei, C-L, Rowe GT, Escobar-Briones E, Boetius A, Soltwedel T, Caley MJ, Soliman Y, Huettmann F, Qu FY, Yu ZS, Pitcher CR, Haedrich RL, Wicksten MK Rex MA, Baguley JG, Sharma J, Danovaro R, MacDonald IR, Nunnally CC, Deming JW, Montagna P, Levesque M, Węsławski JM, Włodarska-Kowalczuk M, Ingole BS, Bett BJ, Billett DSM, Yool A, Bluhm BA, Iken K, Narayanaswamy BE, 2011, Global Patterns and Predictions of Seafloor Biomass Using Random Forests, PLOS ONE 5
  • Węsławski JM, Kendall MA, Włodarska- Kowalczuk M., Iken K, Legeżyńska J., Kedra M, Sejr M. 2011, Climate change effects on Arctic fjord and coastal macrobenthic diversity. Marine Biodiversity 41, 71-85
  • Włodarska-Kowalczuk M., Węsławski J.M., Warzocha J., Janas U. 2010 Habitat loss and possible effects on local species richness in a species-poor system - a case study of southern Baltic Sea macrofauna. Biodiversity & Conservation 19, 3991-4002
  • Sejr MK, Włodarska-Kowalczuk, M, Legeżyńska J, Blicher ME., 2010, Macrobenthic species composition and diversity in the Godthaabsfjord system, SW Greenland. Polar Biology 33, 421-431
  • Kedra M, Włodarska-Kowalczuk M, Węsławski JM. 2010. Decadal change in macrobenthic soft-bottom community structure in a high Arctic fjord (Kongsfjorden, Svalbard). Polar Biology 33, 1-13
  • Kedra M., Gromisz S., Jaskuła R., Legeżyńska J., Maciejewska B., Malec E., Opanowski A., Ostrowska K., Włodarska- Kowalczuk M., Węsławski JM., 2010, Soft bottom fauna of an All Taxa Biodiversity Site - Hornsund (77∨dm;N, Svalbard). Polish Polar Research 31, 309-326
  • Vandepitte L, Vanhoorne B, Kraberg A, Anisimova N, Antoniadou C, Araújo R, Bartsch I, Beker B, Benedetti-Cecchi L, Bertocci I, Cochrane S, Cooper K, Craeymeersch J, Christou E, Crisp DJ, Dahle S, de Boissier M, de Kluijver M, Denisenko S, De Vito D, Duineveld G, Escaravage V, Fleischer D, Fraschetti S, Giangrande A, Heip C, Hummel H, Janas U, Karez R, Kedra M, Kingston P, Kuhlenkamp R, Libes M, Martens P, Mees J, Mieszkowska N, Mudrak S, Munda I, Orfanidis S, Orlando-Bonaca M, Palerud R, Rachor E, Reichert K, Rumohr H, Schiedek D, Schubert P, Sistermans WCH, Sousa Pinto I, Southward AJ, Terlizzi A, Tsiaga E, van Beusekom JEE, Vanden Berghe E, Warzocha J, Wasmund N, Węsławski JM, Widdicombe C, Włodarska-Kowalczuk M, Zettler ML2010 Data integration for European marine biodiversity research: creating a database on benthos and plankton to study large-scale patterns and long-term changes. Hydrobiologia 644, 1-13
  • Węsławski JM, Opanowski A., Legeżyńska J., Maciejewska B., Włodarska- Kowalczuk M, Kędra M., 2010, How many roles can a species play ? Hidden diversity in Arctic crustaceans. Polish Polar Research 31, 205- 216
  • Terlizzi, A., Anderson, M.A., Bevilacqua, S., Fraschetti, S., Włodarska-Kowalczuk, M., Ellingsen, K., 2009, Beta diversity and taxonomic sufficiency do higher-level taxa reflect heterogeneity in species composition? Diversity and Distributions 15, 450-458

    Włodarska-Kowalczuk, M., Kuklinski, P., Ronowicz, M., Legezynska, J., Gromisz, S. 2009. Assessing species richness of macrofauna associated with macroalgae in Arctic kelp forests (Hornsund, Svalbard). Polar Biology 32, 897-905
  • Włodarska-Kowalczuk, M., 2009, Spitsbergen w: Gillespie, R. & Clague, D. (eds.) Encyclopedia of Islands, University of California Press, Berkeley, 865-866
  • Arvanitidis C, Somerfield PJ, Rumohr H, Faulwetter S, Valavanis V, Vasileiadou A, Chatzigeorgiou G, Vanden Berghe E, Vanaverbeke J, Labrune C, Grémare A, Zettler ML, Kedra M, Włodarska-Kowalczuk M, Aleffi IF, Amouroux JM, Anisimova N, Bachelet G, Büntzow M, Cochrane S, Costello MJ, Craeymeersch J, Dahle S, Degraer S, Denisenko S, Dounas C, Duineveld G, Emblow C, Escavarage V, Fabri MC, Fleischer D, Gray JS, Heip C, Hermann M, Hummel H, Janas U, Karakassis I, Kendall MK, Kingston P, Kotwicki L, Laudien J, Mackie ASY, Nevrova HL, Occhipinti-Ambrogi A, Oliver PG, Olsgard F, Palerud R, Petrov A, Rachor E, Revkov NK, Rose A, Sardá R, Sistermans WCH, Speybroeck J, Van Hoey G, Vincx M, Whomersley P, Willems W, Zenetos A. 2009. Biological geography of the European Seas: results from the macrofaunal inventory of the soft-substrate communities, Marine Biology Progress Series, 382: 265-278
  • Escaravage V, Herman PMJ, Merckx B, Włodarska-Kowalczuk M, Amouroux JM, Degraer S, Grémare A, Heip CHR, Hummel H, Karakassis I, Labrune C, Willems W. 2009. Distribution patterns of macrofaunal species diversity in subtidal soft sediments: biodiversity-productivity relationships from the MacroBen database. Marine Ecology Progress Ecology 382: 253-264.
  • Grémare A, Labrune C, Vanden Berghe E, Amouroux JM, Bachelet G, Zettler ML, Vanaverbeke J, Fleischer D, Bigot L, Maire O, Deflandre B, Craeymeersch J, Degraer S, Dounas C, Duineveld G, Heip C, Herrmann M, Hummel H, Karakassis I, Kedra M, Kendall M, Kingston P, Laudien J, Occhipinti A, Rachor E, Sardá R, Speybroeck J, Van Hoey G, Vincx M, Whomersley P, Willems W, Włodarska-Kowalczuk M, Zenetos A. 2009. Comparison of the performances of two biotic indices based on a soft-bottom macrozoobenthos pan-European database, Marine Biology Progress Series, 382: 297-311
  • Renaud PE, Webb TJ, Bjoergesaeter A, Karakassis I, Kedra M, Kendall MA, Labrune C, Lampadariou N, Somerfield PJ, Włodarska-Kowalczuk M, Vanden Berghe E, Claus S, Aleffi IF, Amouroux JM, Bryne KH, Cochrane SJ, Dahle S, Degraer S, Denisenko SG, Deprez T, Dounas C, Fleischer D, Gil J, Gremare A, Janas U, Mackie ASY, Palerud R, Rumohr H, Sarda R, Speybroeck J, Taboada S, van Hoey G, Węsławski JM, Whomersley P, Zettler ML. 2009. Contribution to the Theme Section 'Large-scale studies of the European benthos' Continental-scale patterns in benthic invertebrate diversity: insights from the MacroBen database. Marine Ecology Progress Series 382: 239-252
  • Somerfield PJ, Arvanitidis C, Faulwetter S, Chatzigeorgiou G, Vasileiadou A, Amouroux J, Anisimova N, Cochrane S, Craeymeersch J, Dahle S, Denisenko S, Dounas K, Duineveld G, Grémare A, Heip C, Herrmann M, Karakassis I, Kedra M, Kendall M, Kingston P, Kotwicki L, Labrune C, Laudien J, Nevrova H, Nicolaidou A, Occhipinti-Ambrogi A, Palerud R, Petrov A, Rachor E, Revkov N, Rumohr H, Sardá R, Janas U, Vanden Berghe E, Włodarska-Kowalczuk M, 2009, Assessing evidence for random assembly of marine benthic communities from regional species pools, Marine Biology Progress Series, 382: 279-286
  • Vanden Berghe, E., Claus, S., Appeltans, W., Faulwetter, S., Arvanitidis, C., Somerfield, P. J., Aleffi, I. F., Amouroux, J. M., Anisimova, N., Bachelet, G., Cochrane, S., Costello, M. J., Craeymeersch, J., Dahle, S., Degraer, S., Denisenko, S., Dounas, C., Duineveld, G., Emblow, C., Escaravage, V., Fabri, M. C., Fleischer, D., Grémare, A., Herrmann, M., Hummel, H., Karakassis, I., Kedra, M., Kendall, M., Kingston, P., Kotwicki, L., Labrune, C., Laudien, J., Nevrova, H., Occhipinti, A., Olsgard, F., Palerud, R., Petrov, A., Rachor, E., Revkov, N., Rumohr, H., Sardá, R., Sistermans, W. C. H., Speybroeck, J., Janas, U., Van Hoey, G., Vincx, M., Whomersley, P., Willems, W., Włodarska-Kowalczuk, M., Zenetos, A., Zettler, M. L., Heip C. 2009. "MacroBen" integrated database on benthic invertebrates of European continental shelves: a tool for large-scale analysis across Europe, Marine Biology Progress Series, 382: 225-238
  • Webb, T.J., Aleffi, I.F., Amouroux, J.M, Bachelet, G., Degraer, S, Dounas, C., Fleischer, D., Grémare, A., Herrmann, M., Hummel, H., Karakassis, I., Kedra, M., Kendall, M.A., Kotwicki, L., Labrune, C., Nevrova, E.L., Occhipinti-Ambrogi, A., Petrov, A., Revkov, N.K., Sardá, R., Simboura, N., Speybroeck, J., Van Hoey, G., Vincx, M., Whomersley, P., Willems, W., Włodarska-Kowalczuk, M. 2009. Macroecology of the European soft sediment benthos: insights from the Macroben database, Marine Ecology Progress Series, 382: 287-296
  • Kuklinski, P., Barnes D.K.A., Włodarska-Kowalczuk, M. Gastropods shells, hermit crabs and Arctic bryozoan richness. Bryozoan Research 2007: Proceedings of the 14th International Bryozoology Association Conference, Boone, North Carolina, July 1-8, 2007. Virginia Museum of Natural History Special Publication 2008. no. 15: 93-100.
  • Ronowicz, M., Włodarska-Kowalczuk M., Kuklinski P., 2008, Factors influencing hydroid (Cnidaria, Hydrozoa) biodiversity and distribution in Arctic kelp forests. Journal of the Marine Biological Association of the United Kingdom 88, 1567-1575
  • Kędra M, Włodarska-Kowalczuk M. 2008, Distribution and diversity of sipunculan fauna in high Arctic fjords (west Svalbard), Polar Biology 31, 1181–1190
  • Włodarska-Kowalczuk M., Węsławski, J.M., 2008, Mesoscale spatial structures of soft-bottom macrozoobenthos: physically controlled/impoverished versus biologically accommodated/diverse communities, Mar Ecol Prog Ser 356, 215-224
  • Włodarska-Kowalczuk M., Kedra M., 2007, Surrogacy in natural patterns of benthic distribution and diversity: selected taxa versus lower taxonomic resolution, Mar Ecol Prog Ser , 351, 53-63
  • Barnes D., Kuklinski, P., Włodarska-Kowalczuk M.,2007, Richness, abundance and shell use of subArctic and Arctic hermit crabs, Mar Biol , 152, 1133-1142
  • Włodarska-Kowalczuk M.,Szymelfenig M., Zajączkowski M., 2007, Dynamic sedimentary environments of an Arctic glacier-fed river estuary (Adventfjorden, Svalbard). II. Meio- and macrobenthic fauna, Estuar Coast Shelf Sci, 74, 274-284
  • Zajączkowski M., Włodarska-Kowalczuk M., 2007, Dynamic sedimentary environments of an Arctic glacier-fed river estuary (Adventfjorden, Svalbard). I. Flux, deposition and dynamics of the sediment, Estuar Coast Shelf Sci, 74, 285-296
  • Włodarska-Kowalczuk M., Sicinski, J., Gromisz, S., Kendall, M.A., Dahle, S., 2007, Similar soft-bottom polychaete diversity in Arctic and Antarctic marine inlets, Mar Biol, 151, 607-616
  • Włodarska-Kowalczuk M., 2007, Molluscs in Kongsfjorden (Spitsbergen, Svalbard): a species list and patterns of distribution and diversity, Polar Res, 26, 48-63
  • Renaud, P.E., Włodarska-Kowalczuk M., Trannum H., Holte B., Węsławski J.M., Cochrane S., Dahle S., Gulliksen B., 2007, Multidecadal stability of benthic community struct ure in a high-Arctic glacial fjord (van Mijenfjord, Spitsbergen), Polar Biol, 30, 295-305
  • Włodarska-Kowalczuk, M., Pearson,T.H., Kendall, M.A., 2005, Benthic response to chronic natural physical disturbance by glacial sedimentation in an Arctic fiord , Mar Ecol Prog Ser,303, 31-41
  • Soltwedel, T., Bauerfeind, E., Bergmann, M., Budaeva, N., Hoste, E., Jaeckisch, N., von Juterzenka, K., Matthiessen, J.,Mokievsky, V., Nöthig, E.-M., Quéric, N.-V. Sablotny,B., Sauter, E., Schewe, I., Urban-Malinga, B., Wegner, J., Włodarska-Kowalczuk, M., Klages, M., 2005, HAUSGARTEN - multidisciplinary investigations at a deep-sea long-term observatory in the Arctic Ocean, Oceanography 18, 46-61
  • Kaczmarek, H., Włodarska-Kowalczuk, M., Legezynska, J., Zajączkowski, M., 2005, Shallow sublittoral fauna in Kongsfjord (west Spitsbergen, Svalbard), Polish Polar Res, 26, 137-155
  • Jankowska, K., Włodarska-Kowalczuk, M., Wieczorek, P., 2005, Abundance and biomass of bacteria in two Arctic glacial fjords, Polish Polar Res, 26, 77-84
  • Włodarska-Kowalczuk, M., Kendall, M.A., Węsławski, J.M., Klages, M., Soltwedel, T., 2004, Depth gradients of benthic standing stock and diversity on the continental margin at a high latitude ice-free site (off West Spitsbergen, 79 °N)., Deep-Sea Res I, 51, 1903-1914
  • Włodarska-Kowalczuk, M., Pearson, T., 2004, Soft-bottom macrobenthic faunal associations and factors affecting species distributions in an Arctic glacial fjord (Kongsfjord, Spitsbergen), Polar Biol, 27, 155-167
  • Węsławski, J.M., Włodarska-Kowalczuk, M., Legezynska, J., 2003. Occurrence of soft bottom macrofauna along the depth gradient in High Arctic, 79 °N. Polish Polar Res 24, 73-88.
  • Hop H., Pearson T., Hegseth E.N., Kovacs K.M., Wiencke C., Kwasniewski S., Eiane K., Mehlum F., Gulliksen B., Włodarska-Kowalczuk, M., Lydersen C., Węsławski J.M., Cochrane S., Gabrielsen G.W., Leakey R., Lønne O.J., Zajączkowski M., Falk-Petersen S., Kendall M., Wängberg S.-Å., Bischof K., Voronkov A., Kovaltchouk N.A., Wiktor, J., Poltermann M., di Prisco G., Papucci C., Gerland S. (2002) The marine ecosystem of Kongsfjorden, Svalbard. Polar Res 21, 167-208
  • Rogers, A., Billett, D.S.M., Berger, W., Flach, E., Freiwald, A., Gage, J.D., Hebbeln, D., Heip, C., Pfannkuche, O., Ramirez-Llodra, E.Z., Medlin,L., Sibuet, M., Soetart, K., Tendal, O., Vanreusel, A., Włodarska-Kowalczuk, M., 2002, Life at the edge: achieving prediction from environmental variability and biological variety, In: Wefer, G., Billett, D.S.M. et al. eds. Ocean Margin Systems, Springer, Berlin, 387-404.
  • Węsławski JM, Włodarska-Kowalczuk M, Legezynska J, Kotwicki L 2002 Bioróznorodnosc arktycznego bentosu od szelfu do basenu oceanicznego w Arktyce europejskiej na 79 równolezniku.Przeglad Zoologiczny, XLVI, 3-4, 113-126
  • Włodarska-Kowalczuk, M., Węsławski, J.M., 2001, Impact of climate warming on Arctic benthic biodiversity: a case study of two Arctic glacial bays, Climate Res 18, 127-132.
  • Włodarska-Kowalczuk, M., Szymelfenig M., Kotwicki L, 1999, Macro- and meiobenthic fauna of the Yoldiabukta glacial bay (Isfjorden, Spitsbergen), Polish Polar Res 20, 367-386.
  • Kotwicki L., Włodarska-Kowalczuk, M., Wieczorek P., 1999, Macrozoobenthos of the sandy littoral zone in the military area between Hel and Jurata, Oceanological Studies 28, 97-107.
  • Włodarska-Kowalczuk, M., Węsławski, J.M., Kotwicki, L., 1998, Spitsbergen glacial bays macrofauna - a comparative study, Polar Biology, 20, 66-73
  • Włodarska-Kowalczuk, M., Janas, U., Szaniawska, A., 1996, Hydrogen sulphide and other factors influencing the macrobenthic community structure in the Gulf of Gdansk, Oceanologia, 38, 379-394.
  • Włodarska, M., Węsławski J.M., Gromisz, S., 1996, A comparison of the macrofaunal community structure and diversity in two arctic glacial bays - a ‘cold’ one off Franz Josef Land and a ‘warm’ one off Spitsbergen, Oceanologia, 38, 251-283

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